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Reappraisal of Austrosaurus mckillopi Longman, 1933 from the Allaru Mudstone of Queensland, Australia’s first named Cretaceous sauropod dinosaur

机译:1933年澳大利亚昆士兰州allaru mudstone重新鉴定了澳大利亚第一个名为白垩纪蜥脚类恐龙的南澳龙(Longosa)mmanillopi Longman

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摘要

Austrosaurus mckillopi Longman, 1933 was the first Cretaceous sauropod reported from Australia, and the first Cretaceous dinosaur reported from Queensland (northeast Australia). This sauropod taxon was established on the basis of several fragmentary presacral vertebrae (QM F2316) derived from the uppermost Lower Cretaceous (upper Albian) Allaru Mudstone, at a locality situated 77 km west-northwest of Richmond, Queensland. Prior to its rediscovery in 2014, the type site was considered lost after failed attempts to relocate it in the 1970s. Excavations at the site in 2014 and 2015 led to the recovery of several partial dorsal ribs and fragments of presacral vertebrae, all of which clearly pertained to a single sauropod dinosaur. The discovery of new material belonging toof the type individual of Austrosaurus mckillopi, in tandem with a reassessment of the type seriesmaterial collected in the 1930s, has facilitated the rearticulation of the specimen. The resultant vertebral series comprises six presacral vertebrae—the posteriormost cervical and five anteriormost dorsals—in association with five left dorsal ribs and one right one. The fragmentary nature of the type specimen has historically hindered assessments of the phylogenetic affinities of Austrosaurus, as has the fact that these evaluations were often based on a subset of the type material. The reappraisal of the type series of Austrosaurus presented herein, on the basis of both external morphology and internal morphology visualised through CT data, validates it as a diagnostic titanosauriform taxon (, tentatively placed in Somphospondyli),, and characterised by the possession of an accessory lateral pneumatic foramen on dorsal vertebra I (a feature which appears to be autapomorphic) and by the presence of a robust ventral midline ridge on the centra of dorsal vertebrae I and II. The interpretation of the anteriormost preserved vertebra in Austrosaurus as a posterior cervical has also prompted the re-evaluation of an isolated, partial, posterior cervical vertebra (QM F6142, the “Hughenden sauropod”) derived from the upper Albian Toolebuc Formation (which underlies the Allaru Mudstone). Although this vertebra preserves an apparent unique character of its own (a spinopostzygapophyseal lamina fossa), it is not able to be referred unequivocally to Austrosaurus and is retained as Titanosauriformes indet. Austrosaurus mckillopi is one of the oldest known sauropods from the Australian Cretaceous based on skeletal remains, and potentially provides phylogenetic and/or palaeobiogeographic context for later taxa such as Wintonotitan wattsi, Diamantinasaurus matildae and Savannasaurus elliottorum.
机译:Austrosaurus mckillopi Longman,1933年,是澳大利亚报道的第一只白垩纪蜥脚类恐龙,也是昆士兰州(澳大利亚东北部)报道的第一只白垩纪恐龙。该蜥脚类分类单元是基于几个碎裂的pre前椎骨(QM F2316)建立的,该椎体来源于昆士兰州里士满市西北西北77公里处的最上白垩统(上阿尔比亚)Allaru Mudstone。在2014年重新发现之前,该类型站点在1970年代尝试重新定位失败后被视为丢失。 2014年和2015年在该地点进行的挖掘工作使recovery骨的部分背肋骨和fragments前椎骨的碎片得以恢复,所有这些显然都与单个蜥脚类恐龙有关。在重新评估1930年代收集的系列材料的同时,发现了属于Austrosaurus mckillopi类型个体的新材料,这促进了标本的重新组织。最终的椎骨系列包括六个s前椎骨(最后颈和五个最前背),以及五个左背肋和一个右背肋。类型标本的碎片性质在历史上阻碍了对澳大利亚龙的系统发生亲和力的评估,事实上,这些评估通常基于类型材料的子集。根据通过CT数据可视化的外部形态和内部形态,对本文提出的Austrosaurus类型系列进行重新评估,将其验证为诊断性钛金龙类分类单元(暂时放置在Somphospondyli中),并具有附属物的特征。背椎骨I上有侧向气孔(此特征看来是自发性的),并且在背椎骨I和II的中央存在坚固的腹中线脊。将Austrosaurus中最前保留的椎骨解释为后颈椎,也促使对来自上层Albian Toolebuc地层(其基础是Albian Toolebuc组)的孤立的部分后颈椎骨(QM F6142,“ Hughenden sauropod”)进行重新评估。 Allaru Mudstone)。尽管该椎骨保留了其自身的明显独特特征(脊柱后凸后椎板椎窝),但不能将其明确地称为鼓龙,并保留为泰龙形。澳洲龙mckillopi是澳大利亚白垩纪最古老的蜥脚类动物之一,基于骨骼遗骸,可能为以后的分类群(如Wintonotitan wattsi,Diamantinasaurus matildae和Savannasaurus elliottorum)提供系统发育和/或古生物地理背景。

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